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Beavers are large, semiaquatic rodents in the genus Castor native to the temperate Northern Hemisphere. There are two extant species: the North American beaver (Castor canadensis) and the Eurasian beaver (C. fiber). Beavers are the second-largest living rodents after the capybaras. They have stout bodies with large heads, long chisel-like incisors, brown or gray fur, hand-like front feet, webbed back feet and flat, scaly tails. The Eurasian beaver has a more elongated skull with a more triangular nasal bone opening, lighter fur color and a narrower tail. The animals can be found in a number of freshwater habitats, such as rivers, streams, lakes and ponds. They are herbivorous, consuming tree bark, aquatic plants, grasses and sedges. Beavers build dams and lodges using tree branches, vegetation, rocks and mud; they chew down trees for building material. Dams impound water and lodges serve as shelters. Their infrastructure creates wetlands used by many other species, and because of their effect on other organisms in the ecosystem, they are considered a keystone species. Adult males and females live in monogamous pairs with their offspring. When they are old enough, the young will help their parents repair dams and lodges and may also help raise newly born offspring. Beavers hold territories and mark them using scent mounts made of mud, debris and castoreum, a urine-based substance excreted through the beaver's castor sacs. Beavers can also recognize their kin by their anal gland secretions and are more likely to tolerate them as neighbors. Historically, beavers have been hunted for their fur, meat and castoreum. Castoreum has been used in medicine, perfume and food flavoring, while beaver pelts have been a major driver of the fur trade. Before protections began in the 19th and early 20th centuries, overhunting had nearly exterminated both species. Their populations have rebounded, and they are both listed as least concern by the IUCN Red List of mammals. In human culture, the beaver symbolizes industriousness and is the national animal of Canada.